Lemon Fresh has seen better days.

After the climactic battle in Babel, she finds herself separated from Ezekiel and Cricket in the wastelands. Lemon's abilities to manipulate electricity mark her as a deviate, and deadly corporate operatives are hunting her to use as a weapon in the war between BioMaas Incorporated and Daedelus Technologies. Instead, Lemon finds herself falling in with a group of fellow deviates-a band of teenagers with astonishing abilities, led by an enigmatic figure known as the Major, who may hold the secrets to Lemon's past.

Meanwhile, Cricket find himself in possession of the puritanical Brotherhood, a religious cult set for a head-on collision with the Major and his band. Searching for Lemon, Ezekiel finds a strange ally in an old enemy, and uncovers a plot that may see him reunited with his beloved Ana.

And inside Babel, a remade Eve hatches a plan to bring an end to the world.